Thursday, 17 August 2023

Modern Synthetic Theory by G. L. Stebbins 1971

 The Modern Synthetic Theory

            In the last few decades, numerous researchers have endeavored to elucidate the mechanisms of evolution based on the modern concept of heredity. This theory amalgamates Darwinian evolution with Mendelian genetics, culminating in a comprehensive and unified theory of evolution commonly referred to as Neo-Darwinian theory. This paradigm was pioneered by several eminent evolutionary biologists, including T. Dobzhansky, J.B.S. Haldane, R.A. Fisher, Sewall Wright, G.L. Stebbins, and Ernst Mayr. In 1971, G. L. Stebbins, a renowned Genetics Professor, expounded upon the synthetic theory and identified five fundamental factors intrinsic to the process of organic evolution.

            This theory delineates the evolution of life through genetic modifications within populations, ultimately resulting in the emergence of new species. Key principles underlying the formation of novel species under this theory encompass gene mutations, alterations in chromosome structure and number, genetic recombination, natural selection, and reproductive isolation. Succinct explanations of each factor are provided below:

Gene Mutation

    Mutation refers to the abrupt, heritable alterations transpiring at the genetic level, which consequently manifest as modifications in an organism's phenotypes or external features. Such changes can materialize through the displacement of a gene due to the influence of internal or external factors. The impact of these forces may lead to gene repositioning or its dislodgment, thereby affecting gene expression. Chromosomal mutations can arise from variations in chromosome configuration or count, encompassing phenomena such as deletion, insertion, and inversion. Mutations stemming from these processes can either yield detrimental or advantageous outcomes for the organism. Harmful mutations might precipitate postnatal or ongoing organismal demise. Conversely, beneficial mutations can engender novel advantageous traits, thereby enhancing an organism's suitability for survival. These advantageous mutations may undergo selection through the natural selection process, gradually accumulating incremental changes over time. Consequently, such mutations instigate diversity within a population.

Changes in structure and number of chromosome

    Each species possesses a distinct and consistent arrangement of chromosomes, universally characteristic of its kind. However, the equilibrium of this chromosomal composition can be disrupted by external or internal influences, notably mutations induced by a range of factors such as radiation, chemical agents, UV rays etc. These mutations have the potential to engender alterations in both the numerical count and structural configuration of chromosomes within a species, thereby initiating variations amidst members of the same species. Should these modifications persist and become hereditary across successive generations, an accumulation of changes over time might culminate in the emergence of novel traits, ultimately fostering the genesis of an entirely new species.

Genetic Recombination

            Genetic recombination refers to the amalgamation of genes originating from diverse sources, or the fusion of genetic material derived from male and female individuals of sexually reproducing organisms. This phenomenon presents a significant opportunity for the amalgamation of genetic traits from both parents, thereby contributing to the genetic makeup of the offspring or newly born individual. This process unfolds during sexual reproduction, precisely at the juncture of gamete or reproductive cell formation. The exchange of genetic material transpires between non-sister chromatids during the meiotic cell division, a phenomenon aptly termed "crossing over." This intricate genetic interplay occurring in sexually reproducing organisms fosters a spectrum of genetic variability within a given population. This diversity of genetic traits plays a pivotal role in equipping new individuals with enhanced adaptability to their environment.

Natural selection

            Every living entity is intricately woven into the fabric of nature, drawing sustenance and existence from its nurturing embrace. However, the journey of life is rife with a myriad of challenges that living beings must navigate. Should an organism prove unable to withstand the vicissitudes of natural transformations, it faces the ominous prospect of extirpation from the Earth's tapestry. In the crucible of nature's crucible, survival hinges upon an organism's aptitude to flex and adapt in the face of diverse environmental shifts. Those organisms endowed with superior adaptive prowess are singled out by nature's discerning hand. Through the agency of natural selection, genes undergo a transformative journey from one generation to the next, fostering a predilection for differential modes of reproduction that bestow heightened resilience upon the succeeding lineage. As a result, the ensuing generations emerge as exemplars of fitness, elegantly poised to flourish amidst the ever-changing tableau of their surroundings.

Reproductive Isolation

            Reproductive isolation entails the hindrance of interbreeding within a population owing to the presence of natural barriers. Alternatively, in scenarios where a limited population becomes secluded due to natural forces, the opportunity for interbreeding diminishes, and the exchange of genetic information is curtailed. Over the passage of time, such isolation can facilitate the emergence of novel species within this population, characterized by distinctions from their parental counterparts. This process assumes a pivotal role in driving the tenets of the synthetic theory of evolution, contributing substantially to the overarching narrative of species diversification.


Fig- Outline of modern synthetic theory


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